Ph.D in Biochemistry from the University of Buenos Aires, Argentina, 1966. Postdoctoral training 1967-1971 with Dr. Martin Rodbell at the NIH. Academic appointments at Northwestern University (Associate Professor of Physiology, 1971-1975), Baylor College of Medicine (Professor of Cell Biology, 1975-1994), University of California at Los Angeles (Professor of Anesthesiology, Biological Chemistry and Molecular, Cell and Developmental Biology, 1994-2001). At NIEHS since 2001.
Honors and Awards: CC Bell Research Professorship of Cell Biology (1986-1994), Member of the US National Academy of Sciences (1995), Foreign Member of the Argentine National Academy of Sciences (2004). Foreign Member of the Argentine Academy of Medicine (2006), NIH Distinguished Investigator (2007). Prize "Luis Federico Leloir for International Cooperation in Science, Technology and Innovation" (Argentina, 2011).